Morpheus is a user-friendly and extensible modeling environment for multiscale, multicellular and spatio-temporal systems biology. Morpheus supports the simulation and statistical analysis of (stochastic) spatial cell-based models coupled bi-directionally with ordinary differential equations, reaction-diffusion systems and SBML models. It allows rapid development of multiscale models in biological terms and mathematical expressions rather than programming code. Its graphical user interface supports the entire workflow from model construction and simulation to visualization, archiving and batch processing. For example SBML models can be imported as intra-cellular submodels into existing spatial tissue models. Morpheus is open-source (https://gitlab.com/morpheus.lab/morpheus) and pre-compiled installers are available for Mac/Win/Linux.

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